Mahabharata 12.293.16

ये गुणाः पुरुषस्येह ये च मातृगुणास्तथाअस्थि स्नायु च मज्जा च जानीमः पितृतो द्विज

ye guṇāḥ puruṣasyeha ye ca mātṛguṇāstathāasthi snāyu ca majjā ca jānīmaḥ pitṛto dvija

Our plain-language meaning of this verse is waiting for scholar review.